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Somatotopic layout of nerve cells and terminals in one trunk segment of the human embryo at five weeks. Diagrammatic sections at arm level (right), and of spinal cord (left). Arrows indicate eventual nerve conduction. s = body surface afference; p = proprioceptive afference; v = visceral afference; dm = distal motor; pm = proximal motor: vm = visceral motor. :,' . .. 6w Fig. 4. Human embryos showing form of organs of perception before neuromotor activity. A: Receptor primordia in late-somite embryo.

The SOFS of B is reduced in thickness but has normal lamination. C lacks laminae a-f and has appearance of a tectum totally denervated by enucleation of the eye at day 3 of incubation. l 440 880 1320 D istance in J'm Fig. 9. Same chick as Figure 8. Radioactive tritiated proline injected into the operated eye on day 20. Embryo sacrificed 6 hr later. Radioactive protein was transported down the optic nerve fibers to the tectum. Grain density plots from the autoradiographs show distribution of labeled terminals in the innervated (A and B) and noninnervated areas (C) as well as transversely across the junction region (D).

Regenerated Afferent Connections in Adult Fish and Amphibia Whatever the influence of receptor excitation by patterned stimulation on the segregation of nerve connections in development of embryos or in metamorphosis, in the adults of lower vertebrates the primary projection from retina to optic tectum may end up with its many neural elements precisely specified so that functional connections may regenerate from eye to brain according to a fixed somatotopic 27 NEUROEMBRYOLOGY AND THE DEVELOPMENT OF PERCEPTION 28 COLWYN TREVARTHEN code that is unresponsive to environmental stimulation.

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